Dr. Jason Dunlop

Chitimia‐Dobler, L., Mey, C., Harms, D., Hammel, J., Dunlop, J., Kotthoff, U., Mans, B. (2026). Redescription of Alloceraea cretacea (Acari: Ixodida) with an additional nymphal fossil added to this species: novel insights into the evolution of the Haematobothrion. Parasitology, 153(5), 625-632. DOI: https://doi.org/10.1017/s0031182026101826

Gerbe, A., Bartel, C., Haug, C., Arce, S.I., Dunlop, J.A., Haug, J.T. (2026). An unusual harvestman from Eocene Baltic amber presenting a previously unknown morphology of the pedipalps. Fossil record, 29(1), 1-9. DOI: https://doi.org/10.3897/fr.29.174226

Dunlop, J.A., Friederichs, A. (2026). Mapping the collection: visualising locality data for myriapods in the Museum für Naturkunde Berlin. Zoosystematics and Evolution, 102(1), 1-15. DOI: https://doi.org/10.3897/zse.102.167065

Bartel, C., Mitov, P.G., Dunlop, J.A., Hammel, J.U. (2026). 3D analyses of the first ortholasmatine harvestmen from European Eocene ambers. Acta Palaeontologica Polonica, 71(1), 95-107. DOI: https://doi.org/10.4202/app.01283.2025

Sherwood, D., Cobo, S.A., Dunlop, J.A., Tsavalas, A.T., Geci, D., Lucas, S.M., Brescovit, A.D. (2025). Reidentification of historical specimens assigned to Cupiennius oculifer (Karsch, 1879) (Araneae: Trechaleidae). Zootaxa, 5723(1), 138-142. DOI: https://doi.org/10.11646/zootaxa.5723.1.8

Bicknell, R.D.C., Dunlop, J., Young, A., Lauer, B., Lauer, R., McCoy, V.E. (2025). Unique, dimple-like exoskeletal structures suggest syn-vivo infestations in Late Carboniferous horseshoe crabs. Biology Letters, 21(12), 20250565. DOI: https://doi.org/10.1098/rsbl.2025.0565

Santos, A.A., Dunlop, J.A., Hernández-Orúe, A., Selden, P.A., Diez, J.B., McLoughlin, S. (2025). Trigonotarbids (Arachnida) hidden in plant debris from a Late Pennsylvanian tropical forest at El Bierzo, Castilla y León, Spain. PalZ, 99(4), 567-580. DOI: https://doi.org/10.1007/s12542-025-00745-w

Wu, Z., Dunlop, J., Klimov, P., Mai, H., Peng, A., Liu, Y. (2025). A new whip scorpion (Arachnida: Thelyphonida) with a phoretic mite (Acariformes: Trochometridiidae) from Mid-Cretaceous Kachin amber. BMC Ecology and Evolution, 25. DOI: https://doi.org/10.1186/s12862-025-02392-w

Dunlop, J., Bartel, C. (2025). A new species of fossil Phrynus Lamarck, 1801, from Dominican Republic amber (Amblypygi: Phrynidae). Zootaxa, 5563(1), 64-72. DOI: https://doi.org/10.11646/zootaxa.5563.1.7

Bartel, C., Dunlop, J.A., Wedmann, S. (2024). Iridescent harvestmen (Arachnida: Opiliones: Sclerosomatidae) from the Eocene of Messel, Germany. Palaeobiodiversity and Palaeoenvironments, 104(4), 963-975. DOI: https://doi.org/10.1007/s12549-024-00607-4

Selden, P.A., Dunlop, J.A. (2024). A remarkable spiny arachnid from the Pennsylvanian Mazon Creek Lagerstätte, Illinois. Journal of Paleontology, 98(3), 395-401. DOI: https://doi.org/10.1017/jpa.2024.13

Dunlop, J.A., Garwood, R.J. (2024). A review of fossil scorpion higher systematics. PeerJ, 12, e18557. DOI: https://doi.org/10.7717/peerj.18557

Chitimia-Dobler, L., Handschuh, S., Dunlop, J.A., Pienaar, R., Mans, B.J. (2024). Nuttalliellidae in Burmese amber: implications for tick evolution. Parasitology, 151(9), 891-907. DOI: https://doi.org/10.1017/S0031182024000477

Knecht, R.J., Benner, J.S., Swain, A., Azevedo-Schmidt, L., Cleal, C.J., Labandeira, C.C., Engel, M.S., Dunlop, J.A., Selden, P.A., Eble, C.F., Renczkowski, M.D., Wheeler, D.A., Funderburk, M.M., Emma, S.L., Knoll, A.H., Pierce, N.E. (2024). Early Pennsylvanian Lagerstätte reveals a diverse ecosystem on a subhumid, alluvial fan. Nature Communications, 15(1), 7876. DOI: https://doi.org/10.1038/s41467-024-52181-0

Sherwood, D., Dunlop, J., Sharp, A. (2024). On the identity of Opopaea euphorbicola Strand, 1909 and first records of three other non-native goblin spiders from Ascension Island (Araneae: Oonopidae). Zootaxa, 5437(1), 125-130. DOI: https://doi.org/10.11646/zootaxa.5437.1.9

Dunlop, J.A., Dernov, V.S. (2023). The first trigonotarbid arachnid from Ukraine. Acta Geologica Polonica, 181-187. DOI: https://doi.org/10.24425/agp.2022.143600

Bartel, C., Dunlop, J.A., Giribet, G. (2023). An unexpected diversity of Cyphophthalmi (Arachnida: Opiliones) in Upper Cretaceous Burmese amber. Zootaxa, 5296(3), 421-445. DOI: https://doi.org/10.11646/zootaxa.5296.3.6

Dunlop, J.A., Garwood, R.J. (2023). The status of two fossils assigned to the scorpion genus Palaeophonus and its interpretation as a senior synonym of Allopalaeophonus. Arachnology, 19(6), 940-943. DOI: https://doi.org/10.13156/arac.2023.19.6.940

Garwood, R.J., Dunlop, J.A. (2023). Consensus and conflict in studies of chelicerate fossils and phylogeny. Arachnologische Mitteilungen: Arachnology Letters, 66(1). DOI: https://doi.org/10.30963/aramit6602

Dunlop, J.A., Erdek, M., Bartel, C. (2023). A new species of camel spider (Arachnida: Solifugae) in Baltic amber. Arachnology, 19(4), 772-776. DOI: https://doi.org/10.13156/arac.2023.19.4.772

Dunlop, J.A., Wellman, C.H., Prendini, L., Shear, W.A. (2023). A pectinal tooth with peg sensilla from an Early Devonian scorpion. The Journal of Arachnology, 51(3). DOI: https://doi.org/10.1636/JoA-S-22-024

Cianferoni, F., Dunlop, J.A., Ceccolini, F. (2023). Atypus affinis Eichwald, 1830 in Tuscany. Quaderno di Studi e Notizie di Storia Naturale della Romagna, 57, 145-150. URL: http://www.ssnr.it/57-7.pdf

Garwood, R.J., Dunlop, J.A. (2023). X-ray microtomography of the late Carboniferous whip scorpions (Arachnida, Thelyphonida) Geralinura britannica and Proschizomus petrunkevitchi. Journal of Systematic Palaeontology, 21(1), 2180450. DOI: https://doi.org/10.1080/14772019.2023.2180450

Bartel, C., Dunlop, J.A. (2023). First eupnoid harvestmen (Arachnida: Opiliones: Eupnoi) from mid-Cretaceous Kachin amber, with notes on sexual dimorphism in Halitherses grimaldii (Arachnida: Opiliones: Dyspnoi). Palaeoentomology, 6(3). DOI: https://doi.org/10.11646/palaeoentomology.6.3.11

Šlapeta, J., Halliday, B., Dunlop, J.A., Nachum-Biala, Y., Salant, H., Ghodrati, S., Modrý, D., Harrus, S. (2023). The “southeastern Europe” lineage of the brown dog tick Rhipicephalus sanguineus (sensu lato) identified as Rhipicephalus rutilus Koch, 1844: Comparison with holotype and generation of mitogenome reference from Israel. Current Research in Parasitology & Vector-Borne Diseases, 3, 100118. DOI: https://doi.org/10.1016/j.crpvbd.2023.100118

Dunlop, J.A. (2023). The first Palaeozoic spider (Arachnida: Araneae) from Germany. PalZ, 97(3), 497-504. DOI: https://doi.org/10.1007/s12542-023-00657-7

Wang, H., Lei, X.J., Luo, C.H., Dunlop, J.A. (2023). First jumping spider (Araneae: Salticidae) from mid-Miocene Zhangpu amber. Palaeoworld, 32(4), 716-720. DOI: https://doi.org/10.1016/j.palwor.2022.06.002

Bisong, P.T., Dunlop, J., Madruga, C. (2023). Mammalian type material from Cameroon in the Museum für Naturkunde Berlin. Zoosystematics and Evolution, 99(2), 503-517. DOI: https://doi.org/10.3897/zse.99.110878

Bartel, C., Dunlop, J.A., Sharma, P.P., Selden, P.A., Tarasov, P.E., Ren, D., Shih, C. (2023). Four new Laniatorean harvestmen (Arachnida: Opiliones) from mid-Cretaceous Burmese amber. Palaeoworld, 32(1), 124-135. DOI: https://doi.org/10.1016/j.palwor.2022.06.006

Siyam, M., Dunlop, J.A., Kovařík, F., Mohammad, A. (2023). Additions to the distribution of Sudanese scorpions. Zoosystematics and Evolution, 99(1), 45-53. DOI: https://doi.org/10.3897/zse.99.90875

Chitimia-Dobler, L., Pfeffer, T., Würzinger, F., Handschuh, S., Dunlop, J.A. (2023). New larval records of the extinct hard tick Compluriscutula vetulum (Arachnida: Ixodida) from Burmese amber, with notes on its morphology. Palaeoworld, 33(5), 1327-1335. DOI: https://doi.org/10.1016/j.palwor.2023.10.002

Knecht, R.J., Benner, J.S., Dunlop, J.A., Renczkowski, M.D. (2023). The largest Palaeozoic whip scorpion and the smallest (Arachnida: Uropygi: Thelyphonida); a new species and a new ichnospecies from the Carboniferous of New England, USA. Zoological Journal of the Linnean Society, 200(3), 690-704. DOI: https://doi.org/10.1093/zoolinnean/zlad088

Liu, J., Dunlop, J.A., Steiner, M., Shu, D. (2022). A cambrian fossil from the chengjiang fauna sharing characteristics with gilled lobopodians, opabiniids and radiodonts. Frontiers in Earth Science, 10, Article Number: 861934. DOI: https://doi.org/10.3389/feart.2022.861934

Bartel, C., Derkarabetian, S., Dunlop, J.A. (2022). A new species of Laniatores (Arachnida: Opiliones) from Eocene Baltic amber with notes on the evolution of Insidiatores. Arachnology Letters, 64, 46-51. DOI: https://doi.org/10.30963/aramit6406

Hradská, I., Opluštil, S., Selden, P., Dunlop, J. (2022). A new species of trigonotarbid arachnid from the Pilsen Basin of the Czech Republic. Bulletin of Geosciences, 97(2), 261-268. DOI: https://doi.org/10.3140/bull.geosci.1842

Chitimia-Dobler, L., Mans, B.J., Handschuh, S., Dunlop, J.A. (2022). A remarkable assemblage of ticks from mid-Cretaceous Burmese amber. Parasitology, 149(6), 820-830. DOI: https://doi.org/10.1017/s0031182022000269

Wang, H., Lei, X.J., Luo, C.H., Dunlop, J.A. (2022). First jumping spider (Araneae: Salticidae) from mid-Miocene Zhangpu amber. Palaeoworld. DOI: https://doi.org/10.1016/j.palwor.2022.06.002

Bartel, C., Dunlop, J.A., Sharma, P.P., Selden, P.A., Tarasov, P.E., Ren, D., Shih, C. (2022). Four new Laniatorean harvestmen (Arachnida: Opiliones) from mid-Cretaceous Burmese amber. Palaeoworld. DOI: https://doi.org/10.1016/j.palwor.2022.06.006

Chitimia-Dobler, L., Dunlop, J.A., Pfeffer, T., Würzinger, F., Handschuh, S., Mans, B.J. (2022). Hard ticks in Burmese amber with Australasian affinities. Parasitology, 1-15. DOI: https://doi.org/10.1017/s0031182022001585

Watz, M., Dunlop, J. (2022). Observations on regeneration of the pedipalp and legs of scorpions.. Euscorpius, 345, 1-5. URL: https://mds.marshall.edu/euscorpius/vol2022/iss345/1/

Dunlop, J.A. (2022). Spider Origins: a Palaeontological Perspective. Arachnology, 19(sp1), 182-190. DOI: https://doi.org/10.13156/arac.2022.19.sp1.182

Mitov, P.G., Dunlop, J.A., Bartel, C. (2021). A case of pedipalpal regeneration in a fossil harvestman (Arachnida: Opiliones). Arachnologische Mitteilungen: Arachnology Letters, 61(1), 65-69. DOI: https://doi.org/10.30963/aramit6110

Dunlop, J.A. (2021). Chelicerates as Parasites. In De Baets, K., Huntley, J.W. (Hrsg.), The Evolution and Fossil Record of Parasitism (315-346). Cham: Springer International Publishing. DOI: https://doi.org/10.1007/978-3-030-42484-8_9

Wang, H., Dunlop, J., Gai, Z., Lei, X., Jarzembowski, E.A., Wang, B. (2021). First mixopterid eurypterids (Arthropoda: Chelicerata) from the Lower Silurian of South China. Science Bulletin, 66(22), 2277-2280. DOI: https://doi.org/10.1016/j.scib.2021.07.019

Mitov, P.G., Perkovsky, E.E., Dunlop, J.A. (2021). Harvestmen (Arachnida: Opiliones) in Eocene Rovno amber (Ukraine). Zootaxa, 4984(1), 43-72. DOI: https://doi.org/10.11646/zootaxa.4984.1.6

Scholtz, G., Staude, A., Dunlop, J.A. (2021). Reply to “Points of view in understanding trilobite eyes”. Nature Communications, 12(1), 2084. DOI: https://doi.org/10.1038/s41467-021-22228-7

Braddy, S.J., Dunlop, J.A., Bonsor, J.A. (2021). The Early Devonian eurypterid Leiopterella tetliei from Arctic Canada. Canadian Journal of Earth Sciences, 58(12), 1301-1307. DOI: https://doi.org/10.1139/cjes-2021-0015

Kury, A., Dunlop, J., Mendes, A. (2020). Chapter 8. On the allocation of some Palaeozoic and Tertiary harvestmen.. In WCO-Lite: online world catalogue of harvestmen (Arachnida, Opiliones). Version 1.0 — Checklist of all valid nomina in Opiliones with authors and dates of publication up to 2018.. Self Published. URL: https://pantheon.ufrj.br/bitstream/11422/13031/3/Kury%20et%20al%20(2020)%20WCO-Lite%20(version%201,%20release%200)%20mode%20AAA.pdf

Chitimia-Dobler, L., Dunlop, J. (2020). Cleaning historical tick specimens using an ultrasonic cleaner. Journal of Natural Science Collections, 7, 92-97.

Grau, J., Dunlop, J., Meixner, M., Tappe, D., Gjerde, B. (2020). The complete mitochondrial genome of the pentastomid Linguatula arctica (Pentastomida) from reindeer (Rangifer tarandus) in Northern Norway. Mitochondrial DNA Part B, 5(3), 3438-3439. DOI: https://doi.org/10.1080/23802359.2020.1823255

Haug, J., Azar, D., Ross, A., Szwedo, J., Wang, B., Arillo, A., Baranov, V., Bechteler, J., Beutel, R., Blagoderov, V., Delclòs, X., Dunlop, J., Feldberg, K., Feldmann, R., Foth, C., Fraaije, R., Gehler, A., Harms, D., Hedenäs, L., Hyžný, M., Jagt, J., Jagt-Yazykova, E., Jarzembowski, E., Kerp, H., Khine, P., Kirejtshuk, A., Klug, C., Kopylov, D., Kotthoff, U., Kriwet, J., Mckellar, R., Nel, A., Neumann, C., Nützel, A., Peñalver, E., Perrichot, V., Pint, A., Ragazzi, E., Regalado, L., Reich, M., Rikkinen, J., Sadowski, E., Schmidt, A., Schneider, H., Schram, F., Schweigert, G., Selden, P., Seyfullah, L., Solórzano-Kraemer, M., Stilwell, J., Van Bakel, B., Vega, F., Wang, Y., Xing, L., Haug, C. (2020). Comment on the letter of the Society of Vertebrate Paleontology (SVP) dated April 21, 2020 regarding “Fossils from conflict zones and reproducibility of fossil-based scientific data”: Myanmar amber. PalZ, 94(3), 431-437. DOI: https://doi.org/10.1007/s12542-020-00524-9

Bartel, C., Dunlop, J., Sharma, P., Selden, P., Ren, D., Shih, C. (2020). Laniatorean harvestmen (Arachnida: Opiliones) from mid-Cretaceous Burmese amber. Cretaceous Research, 104703. DOI: https://doi.org/10.1016/j.cretres.2020.104703

Dunlop, J. (2020). Evolution: A Breath of Fresh Air for Eurypterids. Current Biology, 30(21), R1304-R1306. DOI: https://doi.org/10.1016/j.cub.2020.09.052

Scholtz, G., Staude, A., Dunlop, J. (2019). Trilobite compound eyes with crystalline cones and rhabdoms show mandibulate affinities. Nature Communications, 10(2503). DOI: https://doi.org/10.1038/s41467-019-10459-8

Elsaka, M., Mitov, P., Dunlop, J. (2019). New fossil harvestmen (Arachnida: Opiliones) in the HOFFEINS amber collection. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen, 292(2), 155-169. DOI: https://doi.org/10.1127/njgpa/2019/0815

Bartel, C., Dunlop, J. (2019). Two laniatorid harvestmen (Opiliones: Cladonychiidae) from Eocene Baltic amber. Arachnologische Mitteilungen: Arachnology Letters, 58(1), 9-12. DOI: https://doi.org/10.30963/aramit5804

Künzel, N., Dunlop, J., Scholtz, G. (2019). Morphology and evolution of spider book lungs (Araneae). Arthropod Systematics & Phylogeny, 77(2), 267-284. DOI: https://doi.org/10.26049/ASP77-2-2019-05

Müller, S., Dunlop, J., Kotthoff, U., Hammel, J., Harms, D. (2019). The oldest short-tailed whipscorpion (Schizomida): A new genus and species from the Upper Cretaceous amber of northern Myanmar. Cretaceous Research, 106, 104227. DOI: https://doi.org/10.1016/j.cretres.2019.104227

Dunlop, J., Marusik, Y., Vlaskin, A. (2019). Comparing Arachnids in Rovno Amber with the Baltic and Bitterfeld Deposits. Paleontological Journal, 53(10), 1074-1083. DOI: https://doi.org/10.1134/S0031030119100034